Ruby and Sapphire encountered Plusle and Minun on the Abandoned Ship, who were left behind by their original trainer, a Scientist who invented the Scanner. Together they drove away Team Magma, but lost the Scanner to the evil organization during the process. After Kyogre awoke, Plusle and Minun were washed into the sea by the floods and ended up on the Mirage Island where they met Ruby and Sapphire again. Plusle joined Ruby's team and trained under Tate and Liza for double battles. Plusle and Minun later joined the battle at Sootopolis and fired off the finishing electric blow to end Archie and Maxie's evil schemes.